will a Intel P4 2.6ghz system boot with one stick of ddr memory installed or do both memory slots need filling?
Only if you need more memory. How much have you got?
its not p4 its motherboard that controls memory
It will boot with one stick - always (assuming no faults, of course).You would need two (identical) sticks if the motherboard supports dual memory, and you wish to activate it.
That should read dual-channel memory - sorry.
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